Prof. Kathy T. Mullen

Published Papers

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McGill Vision Research, Dept of Ophthalmology, McGill University, Montreal, Quebec, Canada

Use these links to jump to: 1990-1994, 1995-1999, 2000-2004, 2005-2010, 2011


Mullen, K.T., Beaudot, W.H.A. & Ivanov, I.V.  Evidence that global processing does not limit thresholds for RF shape discrimination. Journal of Vision, 11(3): 3, pp1-21, 2011.

Li, X., Mullen, K.T., Thompson, B. & Hess, R.F.  Effective connectivity anomalies in human amblyopia. NeuroImage, 54, 505–516, 2011.


Mullen, K.T. Thompson, B. & Hess, R.F.  Responses of the human visual cortex and LGN to achromatic and chromatic temporal modulation: an fMRI study. Journal of Vision, 10(13): 13, 1–19, 2010.

Garcia-Suarez, L. & Mullen, K.T. Global motion processing in human color vision: a deficit for second order stimuli. Journal of Vision, 10(14): 20, 1–11, 2010. Free access:

Chappell, M. & Mullen, K.T.  The magnocellular visual pathway and the Flash-Lag illusion. Journal of Vision, 10(11): 24, pp1- 10, 2010. (doi: 10.1167/10.11.24) Free access:

Hess, R.F., Thompson, B., Gole, G. & Mullen, K.T.  The amblyopic deficit and its relationship to geniculo-cortical processing streams. Journal of Neurophysiology, 104(1), pp475-483, 2010.

Leh, S.E., Ptito, A., Schonwiesner, M., Chakravarty, M.M. & Mullen, K.T. Blindsight mediated by an S-cone independent collicular pathway: an fMRI study in hemispherectomized subjects. Journal of Cognitive Neuroscience, 22(4),  pp670-682, 2009.

Medina, J.M. & Mullen, K.T.  Cross-orientation masking in human color vision. Journal of Vision, 9(3):20, pp1-16, 2009.
Free access: Journal of Vision:

Hess, R.F., Thompson, B., Gole, G. & Mullen, K.T.  Deficient responses from lateral geniculate nucleus in humans with amblyopia. European Journal of Neuroscience, 29, 1064-1070, 2009. 
Free access:

Mullen, K.T., Dumoulin, S.O. & Hess, R.F.  Color responses of the human lateral geniculate nucleus: Selective amplification of  S-cone signals between the lateral geniculate nucleus and primary visual cortex measured with high-field fMRI.  European Journal of Neuroscience, 28, 1911-1923, 2008. (PDF)

Michna, M. L. & Mullen, K.T.  The contribution of color to global motion. Journal of Vision, 8 (issue 5 article 10), 1-12, 2008.
Free acess: JOV:

Huang, P-C., Mullen, K.T. & Hess, R.F. Colinear facilitation for isoluminant chromatic and luminance stimuli and their facilitation. Journal of Vision, 7, 1-14, 2007 (no. 11, article 6).  URL ISSN 1534-7362.  Free access: JOV:

Medina, J.M. & Mullen, K.T.  Color-luminance interactions in binocular summation. Vision Research, 47, 1120-1128, 2007. (PDF)

Michna, M. L. & Mullen, K.T.  Chromatic mechanisms mediating motion for S-cone isolating stimuli. Vision Research, 47, 1042-1054, 2007. (PDF)

Mullen, K.T., Dumoulin, S.O., McMahon, K.L., de Zubicarary, G.I. & Hess, R.F.  Sensitivity of human retinotopic visual cortex to red-green, blue-yellow and achromatic contrast. European Journal of Neuroscience, 25, 491-502, 2007. (PDF)

Leh, S.E., Mullen, K.T. &  Ptito, A.  Absence of S-cone input in human blindsight following hemispherectomy. European Journal of Neuroscience, 24, 2954-2960, 2006. (PDF)

Sakurai, M. and Mullen, K.T. Cone weights for the two cone opponent systems in peripheral vision and asymmetries of cone contrast sensitivity. Vision Research, 46, 4346-4354, 2006. (PDF)

Beaudot, W.H.A. & Mullen, K.T. Orientation discrimination in human vision: psychophysics and modeling. Vision Research, 46, 26-46, 2006. (PDF)

Beaudot, W.H.A. & Mullen, K.T. Orientation selectivity in luminance and color vision assessed using 2-d bandpass filtered spatial noise. Vision Research, 45, 687-696, 2005. (PDF).

Pitchford, N.J. & Mullen, K.T.  The role of perception, language and preference in the developmental acquisition of basic colour terms.  Journal of Experimental Child Psychology, 90, 275-302, 2005.  (PDF).

Mullen, K.T., Sakurai, M & Chu, W.  Does L/M cone opponency disappear in human periphery? Perception, 34, 951-959, 2005.  (PDF).


Kogan, C.S., Boutet, I., Cornish, K., Zangenehpour, S., Mullen, K.T., Holden, J.J., DerKaloustian, V.M.,  Andermann, E. & Chaudhuri, A.  Differential impact of the FMR1 gene on visual processing in fragile X syndrome. Brain, 127, 591-601, 2004. (PDF).

Mullen, K.T., Yoshizawa, T., & Baker, C.L. Luminance signals mediate the motion of red-green isoluminant gratings: the role of ìtemporal chromatic aberrationî.   Vision Research, 43, 1235-1247, 2003.(PDF)

Beaudot, W.H.A. & Mullen, K.T.  How long-range is contour integration in human color vision.  Visual Neuroscience, 20, 51-64, 2003. (PDF)

Yoshizawa, T., Mullen, K.T. & Baker, C.L. Failure of signed chromatic apparent motion with luminance masking. Vision Research, 43, 751-759, 2003. (PDF)

Pitchford, N.J. & Mullen, K.T. The development of conceptual colour categories in pre-school children: influence of perceptual categorization. Visual Cognition, 10, 51-77, 2003. (PDF) (NB. No color plate in this pdf, see web site for color Figure 1).

Pitchford, N.J. & Mullen, K.T.  The developmental order of color term acquisition in young children. Perception, 31, 1349-1370, 2002. (PDF)

Mullen, K.T. & Kingdom F.A. Differential distribution of red-green and blue-yellow cone opponency across the visual field. Visual Neuroscience, 19, 1-10, 2002. (PDF)

Mullen, K.T. & Beaudot, W.H. The contribution of color vision to a global shape discrimination task. Vision Research, 42, 565-575, 2002. (PDF)

Sankeralli, M.J., Mullen, K.T. & Hine, T.J. Ratio model serves suprathreshold chromatic-luminance discrimination. Journal of the Optical Society of America A 18, 425-435, 2002. (PDF)

Sankeralli, M.J. & Mullen, K.T. Assumptions concerning orthogonality in threshold-scaled versus cone-contrast colour spaces. Vision Research, 41, 53-55, 2001. (PDF)

Hess, R.F., Beaudot, W.H.A. & Mullen, K.T. Dynamics of contour integration. Vision Research, 41, 1023-1037, 2001. (PDF)

Sankeralli, M.J. & Mullen, K.T. Bipolar or rectified chromatic detection mechanisms? Visual Neuroscience, 18, 127-135, 2001. (PDF)

Beaudot, W.H.A. & Mullen, K.T. Processing time of contour integration: the role of color, contrast and curvature. Perception, 30, 833-853, 2001. (PDF)

Sankeralli, M.J., Chen, J.C., Metha, A.B. & Mullen, K.T. Evidence for mild blue-yellow colour vision deficits immediately following fluorescein angiography. Ophthalmic & Physiological Optics, 20, 137-141, 2000. (PDF)

Mullen, K.T., Beaudot, W.H.A. & McIlhagga, W.H. Contour integration in color vision: a common process for the blue-yellow, red-green and luminance mechanisms? Vision Research, 40, 639-655, 2000. (PDF)

Yoshizawa, T., Mullen, K.T. & Baker, C.L. Absence of a chromatic linear motion mechanism in human vision. Vision Research, 40, 1993-2010, 2000. (PDF)

Beaudot, W.H.A. & Mullen, K.T. Role of chromaticity, contrast, and local orientation cues in the perception of numerosity. Perception, 29, 581-600, 2000. (PDF)


Mullen, K.T. & Losada, M.A. The spatial tuning of color and luminance peripheral vision measured with notch filtered noise masking. Vision Research, 39, 721-731, 1999. (PDF)

Mullen, K.T. & Sankeralli, M.J. Evidence for the stochastic independence of the blue-yellow, red-green and luminance detection mechanisms revealed by subthreshold summation. Vision Research, 39, 733-745, 1999. (PDF)

Woodward T.S, Dixon, M.J., Mullen, K.T., Christensen, K.M & Bub, D.N. Investigation of color agnosia: a focus on the analysis of confusions. Neurocase, 5, 95-108, 1999. (PDF)

Sankeralli, M.J. & Mullen, K.T. A ratio model for suprathreshold hue increment detection. Journal of the Optical Society of America A., 16, 2625-2637, 1999. (PDF)

Baker, C.L., Boulton, J.C. & Mullen, K.T. A nonlinear chromatic motion mechanism. Vision Research, 38, 291-302, 1998. (PDF)

Metha, A.B. & Mullen, K.T. Failure of direction discrimination at detection threshold for both fast and slow chromatic motion. Journal of the Optical Society of America A, 15, 2945-2950, 1998. (PDF)

Mullen, K.T., Cropper, S.J. & Losada, M.A. Absence of linear subthreshold summation between red-green and luminance mechanisms over a wide range of spatiotemporal conditions. Vision Research, 37, 1167-1175, 1997. (PDF)

Metha, A.B. & Mullen, K.T. Red-green and achromatic temporal filters: a ratio model predicts contrast dependent speed perception. Journal of the Optical Society of America A, 14, 984-996, 1997. (PDF )

Sankeralli, M.J. & Mullen, K.T. Postreceptoral chromatic detection mechanisms revealed by noise masking in three-dimensional cone contrast space. Journal of the Optical Society of America A. 14, 2633-2646, 1997. (PDF)

Mullen, K.T., Sankeralli, M.J. & Hess, R.F. Color and luminance vision in human amblyopia: Shifts in isoluminance, contrast sensitivity losses and positional deficits. Vision Research, 36, 645-653, 1996. (PDF)

McIlhagga, W.H. & Mullen, K.T. Contour integration with color and luminance contrast. Vision Research, 36, 1265-1279, 1996. (PDF)

Sankeralli, M.J. & Mullen, K.T. Estimation of the L-, M- and S-cone weights of the post-receptoral detection mechanisms. Journal of the Optical Society of America A, 13, 906-915, 1996. (PDF)

Mullen, K.T. & Kingdom F.A. Losses in peripheral color sensitivity predicted from 'hit & miss' post-receptoral cone connections. Vision Research, 36, 1995-2000, 1996. (PDF)

Cropper, S.J., Mullen, K.T. & Badcock, D.R. Motion coherence across different chromatic axes. Vision Research, 36, 2475-2488, 1996. (PDF)

Metha, A.B. & Mullen, K.T. Temporal mechanisms underlying flicker detection and identification for red-green and achromatic stimuli. Journal of the Optical Society of America A, 13, 1969-1980, 1996. (PDF)

Losada, M.A. & Mullen, K.T. Color and luminance spatial tuning estimated by noise masking in the absence of off-frequency looking. Journal of the Optical Society of America A., 12, 250-260, 1995. (PDF)

Kingdom, F.A.A. & Mullen, K.T. Separating colour and luminance information in the visual system. Spatial Vision, 9, 191-219, 1995.


Losada, M.A. & Mullen, K.T. The spatial tuning of chromatic mechanisms identified by simultaneous masking. Vision Research, 34, 331-341, 1994. (PDF)

Mullen, K.T. & Losada, M.A. Evidence for separate pathways for color and luminance detection mechanisms. Journal of the Optical Society of America A., 11, 3136-3151, 1994. (PDF)

Mullen, K.T. & Boulton, J.C. Absence of smooth motion perception in colour vision. Vision Research, 32, pp483-488, 1992. (PDF)

Mullen, K.T. & Boulton, J.C. Interactions between color and luminance contrast in the perception of motion. Ophthalmic and Physiological Optics, 12, 201-205, 1992.

Hess, R.F. & Mullen, K.T.. Suppression of cone signals by rods in human central vision. Ophthalmic and Physiological Optics, 12, 183-188, 1992.

Mullen, K.T. Colour vision as a post receptoral specialization of the central visual field. Vision Research, 31, 119-130, 1991. (PDF)

Anderson S.A., Mullen, K.T. & Hess, R.F. Human peripheral spatial resolution for achromatic and chromatic stimuli: Limits imposed by optical and retinal factors. Journal of Physiology, 442, 47-64, 1991. Use this link:

Boulton J.C and Mullen K.T. A case in which colour vision is no longer motion blind. Clinical Vision Sciences, 5, 175-184, 1990.

Mullen, K.T. and Kulikowski, J.J. Wavelength discrimination at detection threshold. Journal of the Optical Society of America A, 7, 733-742, 1990. (PDF)


Mullen, K.T., Hess, R.F., Sharpe, L.T. and Zrenner, E. The photoreceptors in atypical achromotopsia. J. Physiology, 417, 123-149, 1989.
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Hess, R.F., Mullen, K.T. and Zrenner, E. Human vision with only short wavelength cone receptors: post receptoral properties. J. Physiology, 417, 151-172, 1989.  Use this link:

Mullen, K.T. Spatial influences on colour opponent contributions to pattern detection. Vision Research, 27, 829-839, 1987. (PDF)

Legge, G.E., Mullen, K.T., Woo, G.C. and Campbell, F.W. Tolerance to visual defocus. J. Opt. Soc. Am. A, 4, 851-863 1987. (PDF)

Mullen, K.T. and Plant, G.T. Anomalies in the appearance of colours and of hue discrimination in optic neuritis. Clinical Vision Science, 1, 287-302, 1987. (PDF)

Mullen, K.T. and Plant, G.T. Colour and luminance vision in human optic neuritis. Brain, 109, 1-13, 1986. (PDF)

Mullen, K.T. and Baker, C.L. Jr. A motion aftereffect from an isoluminant stimulus. Vision Research, 25, 685-688, 1985. (PDF)

Mullen, K.T. The contrast sensitivity of human colour vision to red/green and blue/yellow chromatic gratings. J. Physiology, 359, 381-400, 1985.  Download scanned pdf: (PDF) or for better quality use this link: